Scientific Papers
ResearchGate profile

Discovery consists of seeing what everybody has seen and thinking what nobody thought

Albert Szent-Gyorgyi


  • 92. FOUQUET, A., PELOSO, P., RAWIEN JAIRAM, R., LIMA, A. P., MÔNICO, A. T., ERNST, R., KOK, P. J. R. (2022). Back from the deaf: integrative taxonomy revalidates an earless and mute species, Hylodes grandoculis van Lidth de Jeude, 1904, and confirms a new species of Pristimantis Jiménez de la Espada, 1870 (Anura: Craugastoridae) from the eastern Guiana Shield. Organisms Diversity & Evolution. 10.1007/s13127-022-00564-w [link]
  • 91. KOK, P. J. R., LAKING, A., SMITH, C., BERTI, A., MARTEL, A. & PASMANS, F. 2022. Tourism may threaten wildlife disease refugia. Conservation Letters, e12902. 10.1111/conl.12902 [link]
  • 90. MORAES, L. J. C. L., WERNECK, F. P., RÉJAUD, A., RODRIGUES, M. T., PRATES, I., GLAW, F., KOK, P. J. R., RON, S. R., CHAPARRO, J. C., OSORNO-MUÑOZ, M., DAL VECHIO, F., RECODER, R. S., MARQUES-SOUZA, S., ROJAS, R. R., DEMAY, L., HRBEK, T. & FOUQUET, A. 2022. Diversification of tiny toads (Bufonidae: Amazophrynella) sheds light on ancient landscape dynamism in Amazonia. Biological Journal of the Linnean Society. 10.1093/biolinnean/blac006 [link]
  • 89. FOUQUET, A., CORNUAULT, J., RODRIGUES, M. T., WERNECK, F. P., HRBEK, T., ACOSTA-GALVIS, A. R., MASSEMIN, D., KOK, P. J. R. & ERNST, R. 2022. Diversity, biogeography, and reproductive evolution in the genus Pipa (Amphibia: Anura: Pipidae). Molecular Phylogenetics and Evolution 10.1016/j.ympev.2022.107442 [link]


  • 88KOK, P. J. R., BROHOLM, T. L. & MEBS, D. 2021. Thriving in a hostile world: insights from the dietary strategy of two allopatric, closely-related tepui summit endemic amphibians. Ecology and Evolution, 10.1002/ece3.7682 [link]
  • 87. FOUQUET, A. LEBLANC, K., FABRE, A.-C, RODRIGUES, M. T., MENIN, M., COURTOIS, E. A., DEWYNTER, M., HÖLTING  M., ERNST, R., PELOSO, P. L. V., KOK, P. J. R. 2021. Comparative osteology of the fossorial frogs of the genus Synapturanus (Anura, Microhylidae) with the description of three new species from the Eastern Guiana Shield. Zoologischer Anzeiger, 293(e4941) 10.1016/j.jcz.2021.05.003 [link]
  • 86. FOUQUET, A. LEBLANC, K., FRAMIT, M., RÉJAUD  A., RODRIGUES, M. T., CASTROVIEJO-FISHER, S., PELOSO, P. L. V., PRATES, I., MANZI, S., SUESCUN, U., BARONI, S., MORAES, L., RECODER, R., MARQUES DE SOUZA, S., DAL VECCHIO, F., CAMACHO, A., GUELLERE, J. M., ROJAS-RUNJAIC, F. J. M., GAGLIARDI-URRUTIA, G., DE CARVALHO, V. T., GORDO, M., MENIN, M., KOK, P. J. R., HRBEK, T., WERNECK, F. P., CRAWFORD, A. J., RON, S. R., MUESES-CISNEROS, J. J., ROJAS ZAMORA, R. R., PAVAN, D., SIMÕES, P. I., ERNST, R. & FABRE, A.-C. 2021. Species diversity and biogeography of an ancient frog clade from the Guiana Shield (Anura: Microhylidae: Adelastes, Otophryne, Synapturanus) exhibiting spectacular phenotypic diversification. Biological Journal of the Linnean Society 132(2): 233–256 [link]


  • 85. RÉJAUD, A., RODRIGUES, M. T., CRAWFORD, A. J., CASTROVIEJO-FISHER, S., JARAMILLO, A. F., CHAPARRO, J. C., GLAW, F., GAGLIARDI-URRUTIA, G. L. A., MORAVEC, J., DE LA RIVA, I. J., PEREZ, P., LIMA, A. P., WERNECK, F. P., HRBEK, T., RON, S. R., ERNST, R., KOK, P. J. R., DRISKELL, A., CHAVE, J. & FOUQUET, A. 2020. Historical biogeography identifies a possible role of Miocene wetlands in the diversification of the Amazonian rocket frogs (Aromobatidae: Allobates). Journal of Biogeography 10.1111/jbi.13937 [link]
  • 84. FOUQUET, A., JAIRAM, R., OUBOTER, P. & KOK, P. J. R. 2020. Two new species of Anomaloglossus (Anura: Aromobatidae) of the stepheni group from Suriname. Zootaxa, 4820 (1): 147–164 [link]
  • 83. VACHER, J-P., CHAVE, J., FICETOLA, F., SOMMERIA-KLEIN, G., TAO, S., THÉBAUD, C., BLANC, M., CAMACHO, A., CASSIMIRO, J., COLSTON, T. J., DEWYNTER, M. ERNST, R., GAUCHER, P., OLIVEIRA GOMES, J., JAIRAM, R., KOK, P. J. R., DIAS LIMA, J., MARTINEZ, Q., MARTY, C., NOONAN, B. P., NUNES, P., OUBOTER, P., RECODER, R., RODRIGUES, M. T., SNYDER, A. MAQUES-SOUZA, S. & FOUQUET, A. 2020. Large scale DNA-based survey of frogs in Amazonia suggests a vast underestimation of species richness and endemism. Journal of Biogeography 10.1111/jbi.13847 [link]
  • 82. KOK, P. J. R., VAN DER VELDEN, M., MEANS, D. B., RATZ, S., JOSIPOVIC, I., BOONE, M. & MCDIARMID, R. 2020. Coping with the extremes: comparative osteology of the tepui-associated toad Oreophrynella and its bearing on the evolution of osteological novelties in the genus. Zoological Journal of the Linnean Society 10.1093/zoolinnean/zlz172 [link]


  • 81KOK, P. J. R, VAN DOORN, L. & DEZFOULIAN, R. 2019. Predation by non-bioluminescent firefly larvae on a tepui-summit endemic toad. Current Biology29(22): R1170-R1171 [link]
  • 80. FOUQUET, A., VACHER, J.-P., COURTOIS, E. A., DESCHAMPS, C., OUBOTER, P., JAIRAM, R., GAUCHER, P., DUBOIS, A &, KOK, P. J. R. 2019. A new species of Anomaloglossus (Anura: Aromobatidae) of the stepheni group with the redescription of A. baeobatrachus (Boistel and de Massary, 1999), and an amended definition of A. leopardus Ouboter and Jairam, 2012. Zootaxa, 4576: 439-460 [link]
  • 79. TORRES-CARVAJAL, O., ECHEVARRIA, L., LOBOS, S. E., VENEGAS, P. J. & KOK, P. J. R. 2019. Phylogeny, diversity and biogeography of Neotropical sipo snakes (Serpentes: Colubrinae: Chironius). Molecular Phylogenetics and Evolution, 130: 315-329 [link]
  • 78. PINHEIRO, P. D. P., KOK, P. J. R., NOONAN, B. P., MEANS, D. B., HADDAD, C. F. B. & FAIVOVICH, J. 2019. A new genus of Cophomantini, with comments on the taxonomic status of Boana liliae (Anura: Hylidae). Zoological Journal of the Linnean Society, 185: 226-245 [link]


  • 77KOK, P. J. R., NICOLAÏ, M. P. J., LATHROP, A. & MACCULLOCH, R. D. 2018. Anomaloglossus meansi sp. n., a new Pantepui species of the Anomaloglossus beebei group (Anura, Aromobatidae). Zookeys, 759: 99-116 [link]
  • 76. MARQUES-SOUZA, S., PRATES, I., FOUQUET, A., CAMACHO, A., KOK, P. J. R., NUNES, P. M. S., DAL VECHIO, F., SOUSA RECODER, R., MEJIA, N., TEIXEIRA JUNIOR, M., BARRIO-AMORÓS, C., CASSIMIRO, J., DIAS LIMA, J., DE SENA, M. A. & RODRIGUES, M. T. 2018. Reconquering the water: evolution and systematics of South and Central American aquatic lizards (Gymnophthalmidae). Zoologica Scripta, 47: 255-265 [link]
  • 75KOK, P. J. R., BITTENBINDER, M. A., VAN DEN BERG, J. K., MARQUES-SOUZA, S., NUNES, P. M. S., LAKING, A., TEIXEIRA JUNIOR, M., FOUQUET, A., MEANS D. B., MACCULLOCH, R. D. & RODRIGUES, M. T. 2018. Integrative taxonomy of the gymnophthalmid lizard Neusticurus rudis Boulenger, 1900 identifies a new species in the eastern Pantepui region, north-eastern South America. Journal of Natural History, 52(13): 1029-1066 [link]
  • 74. FOUQUET A., VACHER J.-P., COURTOIS, E., VILLETTE, B., REIZINE, H., GAUCHER, P., JAIRAM, R., OUBOTER, P. & KOK, P. J. R. 2018. On the brink of extinction: two new species of Anomaloglossus from French Guiana and amended definition of Anomaloglossus degranvillei and A. surinamensis (Anura: Aromobatidae). Zootaxa, 4379: 001-023 [link]
  • 73KOK, P. J. R., DEZFOULIAN R., MEANS D. B., FOUQUET A. & BARRIO-AMORÓS C. L. 2018. Amended diagnosis and new description of Pristimantis marmoratus (Boulenger, 1900) (Amphibia: Craugastoridae), with description of its advertisement call and notes on its breeding ecology and phylogenetic relationships. European Journal of Taxonomy, 397: 1-30 [link]
  • 72KOK, P. J. R., RATZ, S., MACCULLOCH, R. D., LATHROP, A., DEZFOULIAN R., AUBRET, F. & MEANS D. B. 2017. Historical biogeography of the palaeoendemic toad genus Oreophrynella (Amphibia: Bufonidae) sheds a new light on the origin of the Pantepui endemic terrestrial biota. Journal of Biogeography, 45: 26-36 [link]


  • 71. AUBRET F., BIGNON F., HALLE A., BLANVILLAIN G., KOK, P. J. R. & SOUCHET J. 2017. Yolk removal generates hatching asynchrony in snake eggs. Scientific Reports 7, 3041; doi: 10.1038/s41598-017-03355-y [link]
  • 70. VACHER J.-P., KOK, P. J. R., RODRIGUES M. T., DIAS LIMA J., MORENZINI A., MARTINEZ Q., FALLET M., COURTOIS E., BLANC M., GAUCHER P., DEWYNTER M., JAIRAM R., OUBOTER P., THEBAUD C, FOUQUET A. 2017. Cryptic diversity in Amazonian frogs: Integrative taxonomy of the genus Anomaloglossus (Amphibia: Anura: Aromobatidae) reveals a unique case of diversification within the Guiana Shield. Molecular Phylogenetics and Evolution, 112: 158-173 [link]
  • 69. JABLONSKI, D., GRUL’A, D., BARRIO-AMORÓS, C. L. & KOK, P. J. R. 2017. Molecular phylogenetic relationships among Pristimantis summit populations in the eastern tepui chain: insights from P. aureoventris (Anura: Craugastoridae). Salamandra, 53(3): 473-478 [link]
  • 68. KÖHLER, J., JANSEN, M., RODRÍGUEZ, A., KOK, P. J. R., TOLEDO, L. F., EMMRICH, M., GLAW, F., HADDAD, C. F. B., RÖDEL, M.-O. & VENCES, M. 2017. The use of bioacoustics in anuran taxonomy: theory, terminology, methods and recommendations for best practice. Zootaxa, 4251: 1-124 [link]
  • 67. NICOLAI, M. P. J., PORCHETTA, S., BALAKRISHNA, S., BOTHA, D. P. & KOK, P. J. R. 2017. Preliminary observations on the circadian variation in site fidelity in Atelopus hoogmoedi (Lescure, 1974) (Anura, Bufonidae). Amphibian and Reptile Conservation 11(1): 45–50 [link]


  • 66. AUBRET F., BIGNON F., KOK, P. J. R. & BLANVILLAIN G. 2016. Only child syndrome in snakes: Eggs incubated alone produce asocial individuals. Scientific Reports 6, 35752; doi: 10.1038/srep35752 [link]
  • 65KOK, P. J. R., RUSSO, V. G., RATZ, S., MEANS, D. B., MACCULLOCH, R. D., LATHROP, A., AUBRET, F. & BOSSUYT, F. 2016. Evolution in the South American “Lost World”: insights from multilocus phylogeography of stefanias (Anura, Hemiphractidae, Stefania). Journal of Biogeography, DOI: 10.1111/jbi.12860 [link]
  • 64KOK, P. J. R., RUSSO, V. G., RATZ, S., & AUBRET, F. 2016. On the distribution and conservation of two “Lost World” tepui summit endemic frogs, Stefania ginesi Rivero, 1968 and S. satelles Señaris, Ayarzagüena, and Gorzula, 1997. Amphibian and Reptile Conservation 10(1): 5–12 (e115) [link]
  • 63. AUBRET, F., BLANVILLAIN, G., BIGNON, F. & KOK, P. J. R. 2016. Heartbeat, embryo communication and hatching synchrony in snake eggs. Scientific Reports 6: 23519|DOI:10.1038/srep23519 [link]
  • 62. FOUQUET, A., MARTINEZ, Q., ZEIDLER, L., COURTOIS, E. A., GAUCHER, P., BLANC, M., LIMA, J. D., MARQUES SOUZA, S., RODRIGUES, M. T. & KOK, P. J. R. 2016. Cryptic diversity in the Hypsiboas semilineatus species group (Amphibia, Anura) with the description of a new species from the eastern Guiana Shield. Zootaxa 4084(1): 79-104 [link]


  • 61. KOK, P. J. R., RATZ, S., TEGELAAR, M., AUBRET, F. & MEANS, D. B. 2015. Out of taxonomic limbo: a name for the species of Tepuihyla (Anura: Hylidae) from the Chimantá Massif, Pantepui region, northern South America. Salamandra 51(4): 283-314 [link]
  • 60. AUBRET, F., BLANVILLAIN, G. & KOK, P. J. R. 2015.  Myth busting? Effects of embryo positioning and egg turning on hatching success in the water snake Natrix maura. Scientific Reports 5:13385|DOI:10.1038/srep13385 [link]
  • 59. KOK, P. J. R. 2015. A new species of the Pantepui endemic genus Riolama (Squamata: Gymnophthalmidae) from the summit of Murisipán-tepui, with the erection of a new gymnophthalmid subfamily. Zoological Journal of the Linnean Society 174: 500-518 [full pdf on request]
  • 58. FOUQUET, A., MARQUES SOUZA, S., SALES NUNES, P. M., KOK, P. J. R., CURCIO, F. F., DE CARVALHO, C.M., GRANT, T. & RODRIGUES, M. T. 2015. Two new endangered species of Anomaloglossus (Anura: Aromobatidae) from Roraima state, northern Brazil. Zootaxa 3926(2): 191-210 [link]
  • 57. VAN BOCXLAER, I., TREER, D., MAEX, M., VANDEBERGH, W., JANSSENSWILLEN, S., STEGEN, G., KOK, P. J. R., WILLAERT, B., MATTHIJS, S., MARTENS, E., MORTIER, A., DE GREVE, H., PROOST, P. & BOSSUYT, F. 2015. Side-by-side secretion of late Palaeozoic diverged courtship pheromones in an aquatic salamander. Proceedings of the Royal Society B: Biological Sciences DOI: 10.1098/rspb.2014.2960 [link]
  • 56. LEITE, Y. L. R., KOK, P. J. R. & WEKSLER, M. 2015. Evolutionary affinities of the “Lost World” mouse suggest late Pliocene connection between the Guiana and the Brazilian shields. Journal of Biogeography DOI: 10.1111/jbi.12461 [link]


  • 55. GEHARA, M., CRAWFORD, A. J., ORRICO, V. G. D., RODRÍGUEZ, A., LÖTTERS, S., FOUQUET, A., BALDO, D., BARRIENTOS, L. S., BRUSQUETTI, F., CASTROVIEJO-FISHER, S., DE LA RIVA, I., ERNST, R., FAIVOVICH, J. GAGLIARDI URRUTIA, G., GLAW, F., GUAYASAMIN, J., HÖLTING, M., JANSEN, M., KOK, P. J. R., KWET, A., LINGNAU, R., LYRA, M., MORAVEC, J., PADIAL, J. M., POMBAL JR., J., ROJAS-RUNJAIC, F. J. M., SCHULZE, A. SEÑARIS, J. C., SOLÉ, M., RODRIGUES, M. T., TWOMEY, E., HADDAD, C. F. B., VENCES, M. & KÖHLER, J. 2014. High levels of diversity uncovered in a widespread nominal taxon: continental phylogeography of the Neotropical tree frog Dendropsophus minutusPLOS ONE 9(9): 1-12 [link]
  • 54. WILKINSON, M., KOK, P. J. R., AHMED, F. & GOWER, D. J. 2014. The “lungless” Caecilita is a lunged Microcaecilia (Amphibia: Gymnophiona: Siphonopidae): implications for understanding the evolution of lunglessness. Zootaxa 3779(3): 383-388 [link]


  • 53. FOUQUET, A., MARTINEZ, Q., COURTOIS, E. A., DEWYNTER, M., PINEAU, K., GAUCHER, P., BLANC, M., MARTY, C. & KOK, P. J. R. 2013. A new species of the genus Pristimantis (Amphibia, Craugastoridae) associated with the moderately elevated massifs of French Guiana. Zootaxa 3750(1): 569-586 [link]
  • 52. PASSOS, P., KOK, P. J. R., DE ALBUQUERQUE, N. R. & RIVAS, G.  2013. Groundsnakes of the Lost World: A Review of Atractus (Serpentes: Dipsadidae) from the Pantepui region, northern South America. Herpetological Monographs 27: 52-86 [link]
  • 51KOK, P. J. R. 2013. Two new charismatic Pristimantis species (Anura: Craugastoridae) from the tepuis of “The Lost World” (Pantepui region, South America). European Journal of Taxonomy 60: 1-24 [link]
  • 50. ROELANTS, K., FRY, B. G., YE, L., NORMAN, J. A., STIJLEMANS, B., KOK, P. J. R., CLYNEN, E., SCHOOFS, L., CORNELIS, P. & BOSSUYT, F. 2013. Origin and functional diversification of an amphibian defense peptide arsenal. PLOS Genetics 9(8): e1003662. doi:10.1371/journal.pgen.1003662 [link]
  • 49KOK, P. J. R. & BARRIO-AMORÓS, C. L. 2013. On the taxonomic validity of Pristimantis tepuiensis (Schlüter & Rödder, 2007) and P. stegolepis (Schlüter & Rödder, 2007), with remarks on the type series of P. guaiquinimensis (Schlüter & Rödder, 2007). Zootaxa 3694(1): 75-80 [link]
  • 48KOK, P. J. R., HÖLTING, M. & ERNST, R. 2013. A third microendemic to the Iwokrama Mountains of central Guyana: a new “cryptic” species of Allobates Zimmerman and Zimmerman, 1988 (Anura: Aromobatidae). Organisms Diversity and Evolution DOI: 10.1007/s13127-013-0144-4 [link]
  • 47KOK, P. J. R., MEANS, D. B. & RIVAS, G. A. 2013. First record of the genus Anadia (Reptilia, Squamata, Gymnophthalmidae) from Guyana based on an enigmatic specimen from Mount Kopinang, Wokomung Massif. Boletim do Museu Paraense Emílio Goeldi 8(1): 27-39 [pdf]
  • 46. JUNGFER, K-H, FAIVOVICH, J., PADIAL, J. M., CASTROVIEJO-FISHER, S., LYRA, M., VON MULLER BERNECK, B., IGLESIAS, P., KOK, P. J. R., MACCULLOCH, R. D., RODRIGUES, M. T., VERDADE, V. K., TORRES GASTELLO, C. P., CHAPARRO, J. C., VALDUJO, P. H., REICHLE, S., MORAVEC, J., GVOZDIK, V., GAGLIARDI-URRUTIA, G., ERNST, R., DE LA RIVA, I., MEANS, D. B., LIMA, A. P., SENARIS, J. C., WHEELER, W. & HADDAD, C. F. B. 2013. Systematics of spiny-backed treefrogs (Hylidae: Osteocephalus): an Amazonian puzzle. Zoologica Scripta DOI: 10.1111/zsc.12015 [link]
  • 45.  MEILINK, W. R. M., CLEGG, J. R., MAYERL, C. J., SABINO PINTO, J., GRASSO, D., STEGENS, G., SEGAL, M. & KOK, P. J. R. 2013. Confirmation of the presence of the sphaerodactylid lizard Gonatodes vittatus (Lichtenstein, 1856) in Guyana, and indication of a reproductively active population in Georgetown. Salamandra 49(1): 59-62 [pdf]
  • 44.  KOK, P. J. R., WILLAERT, B. & MEANS, D. B. 2013. A new diagnosis and description of Anomaloglossus roraima (La Marca, 1998) (Anura: Aromobatidae: Anomaloglossinae), with description of its tadpole and call. South American Journal of Herpetology 8(1): 29-45 [link]


  • 43.  FOUQUET, A., LOEBMANN, D., CASTROVIEJO-FISHER, S., PADIAL, J. M., DILL ORRICO, V. G., LYRA, M., JOVENTINO ROBERTO, I., KOK, P. J. R., HADDAD, C. F. B. & RODRIGUES, M. T. 2012. From Amazonia to the Atlantic forest: Molecular phylogeny of Phyzelaphryninae frogs reveals unexpected diversity and a striking biogeographic pattern emphasizing conservation challenges. Molecular Phylogenetics and Evolution 65: 547-561 [link]
  • 42.  SALES NUNES, P. M., FOUQUET, A., CURCIO, F. F., KOK, P. J. R. & RODRIGUES, M. T. 2012. Cryptic species in Iphisa elegans Gray, 1851 (Squamata: Gymnophthalmidae) revealed by hemipenial morphology and molecular data. Zoological Journal of the Linnean Society 166: 361-376 [link]
  • 41.  KOK, P. J. R, MACCULLOCH, R. D., MEANS, D. B., ROELANTS, K., VAN BOCXLAER, I. & BOSSUYT, F. 2012. Low genetic diversity in tepui summit vertebrates. Current Biology 22(15): 589-590 [pdf] Supplementary information [pdf]. Featured as Editor’s Choice in Science 337: 1020 (2012)
  • 40.  RIVAS, G. A., UGUETO, G., MOLINA, C. R., BARROS, T. R., BARRIO-AMORÓS, C. L. & KOK, P. J. R. 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1-64 [link]


  • 39.  KOK, P. J. R. & RIVAS, G. A. 2011. A new species of Anadia (Reptilia, Squamata) from the Venezuelan “Lost World”, northern South America. European Journal of Taxonomy 3: 1-18 [pdf]
  • 38.  KOK, P. J. R. 2011. A new species of the genus Gonatodes Fitzinger, 1843 (Reptilia: Sphaerodactylidae) from central Guyana, northern South America. Zootaxa 3018: 1-12 [link]
  • 37.  KOK, P. J. R, MEANS, D. B. & BOSSUYT, F. 2011. A new highland species of Pristimantis Jiménez de la Espada, 1871 (Anura: Strabomantidae) from the Pantepui region, northern South America. Zootaxa 2934: 1-19 [link]


  • 36.  WILKINSON, M. & KOK, P. J. R. 2010. A new species of Microcaecilia (Amphibia: Gymnophiona: Caeciliidae) from Guyana. Zootaxa 2719: 35-40 [link]
  • 35.  KOK, P. J. R, MACCULLOCH, R. D., LATHROP, A., WILLAERT, B. & BOSSUYT, F. 2010. A new species of Anomaloglossus(Anura: Aromobatidae) from the Pakaraima Mountains of Guyana. Zootaxa 2660: 18-32 [link]
  • 34.  KOK, P. J. R. 2010. A new species of Chironius Fitzinger, 1826 (Squamata: Colubridae) from the Pantepui region, northeastern South America. Zootaxa 2611: 31-44 [link]
  • 33.  DESAMORE, A., VANDERPOORTEN, A., LAENEN, B., GRADSTEIN S. R. & KOK, P. J. R. 2010. Biogeography of the Lost World (Pantepui region, northeastern South America): insights from bryophytes. Phytotaxa 9: 254-265 [pdf]
  • 32.  KOK, P. J. R. 2010. A redescription of Anomaloglossus praderioi (La Marca, 1998) (Anura: Aromobatidae: Anomaloglossinae), with description of its tadpole and call. Papéis Avulsos de Zoologia, 50(4): 51-68 [pdf]
  • 31.  GOWER, D. J., WILKINSON, M., SHERRATT, E. & KOK, P. J. R. 2010. A new species of Rhinatrema Duméril & Bibron (Amphibia: Gymnophiona: Rhinatrematidae) from Guyana. Zootaxa, 2391: 47-60 [pdf]


  • 30.  MACCULLOCH, R. D., LATHROP, A., KOK, P. J. R., ERNST, R. & KALAMANDEEN, M. 2009. The genus Oxyrhopus(Serpentes: Dipsadidae: Xenodontinae) in Guyana: morphology, distributions and comments on taxonomy. Papéis Avulsos de Zoologia, 49(36): 485-495 [pdf]
  • 29.  KOK, P. J. R. 2009. A new species of Oreophrynella (Anura: Bufonidae) from the Pantepui region of Guyana, with notes on Omacconnelli Boulenger, 1900. Zootaxa, 2071: 35-49 [link]
  • 28.  KOK, P. J. R. 2009. Lizard in the clouds: a new highland genus and species of Gymnophthalmidae (Reptilia: Squamata) from Maringma tepui, western Guyana. Zootaxa, 1992: 53-67 [link]
  • 27.  CASTROVIEJO-FISHER, S., GUAYASAMIN, J. M. & KOK, P. J. R. 2009. Species status of Centrolene lema Duellman and Señaris, 2003 (Amphibia: Centrolenidae) revealed by Integrative Taxonomy. Zootaxa, 1980: 16-28 [link]


  • 26.  KOK, P. J. R. & RIVAS FUENMAYOR, G. A. 2008. Typhlophis ayarzaguenai Señaris, 1998 is a junior synonym of Typhlophis squamosus (Schlegel, 1839). Amphibia-Reptilia, 29(4): 555-558 [pdf]
  • 25.  KOK, P. J. R. 2008. A new highland species of Arthrosaura Boulenger, 1885 (Squamata: Gymnophthalmidae) from Maringma tepui on the border of Guyana and Brazil. Zootaxa, 1909: 1-15 [link]
  • 24.  MACCULLOCH, R. D., LATHROP, A., KOK, P. J. R., MINTER, L. R., KHAN, S. Z. & BARRIO-AMORÓS, C. L. 2008. A New Species of Adelophryne (Anura: Eleutherodactylidae) from Guyana, with Additional Description of A. gutturosa. Zootaxa, 1884: 36-50 [link]
  • 23.  KOK, P. J. R. & CASTROVIEJO-FISHER, S. 2008. Glassfrogs (Anura: Centrolenidae) of Kaieteur National Park, Guyana, with notes on the distribution and taxonomy of some species of the family in the Guiana Shield. Zootaxa, 1680: 25-53 [link]


  • 22.  KOK, P. J. R., RIVAS FUENMAYOR, G. A. & PAUWELS, O. S. G. 2007. The taxonomic status of the Venezuelan snakes Atractus matthewi and A. nororientalis (Serpentes, Colubridae). Zootaxa, 1493: 66-68 [link]
  • 21.  KOK, P. J. R., ERNST, R. 2007. A new species of Allobates (Anura: Aromobatidae: Allobatinae) exhibiting a novel reproductive behaviour. Zootaxa, 1555: 21-38 [link]
  • 20.  KOK, P. J. R., KOKUBUM, M. N. C., MACCULLOCH, R. D. & LATHROP, A. 2007. Morphological variation in Leptodactylus lutzi(Heyer, 1975) (Anura: Leptodactylidae) with description of its advertisement call and notes on its courtship behavior. Phyllomedusa, 6(1): 45-60 [pdf]
  • 19.  KOK, P. J. R. & BENJAMIN, P. 2007. Stefania evansi (Groete Creek Carrying Frog): Froglet carrying. The Herpetological Bulletin, 100: 38-39 [pdf]
  • 18.  KOK, P. J. R., BENJAMIN, P. & LENGLET, G. L. 2007. Stefania woodleyi (Woodley’s Treefrog). Defensive behavior. Herpetological Review, 38(2): 198 [pdf]


  • 17.  KOK, P. J. R. 2006. A new species of Hypsiboas (Amphibia: Anura: Hylidae) from Kaieteur National Park, eastern edge of  the Pakaraima Mountains, Guyana. Bulletin de l’Institut Royal des Sciences naturelles de BelgiqueBiologie, 76: 191-200 [pdf]
  • 16.  KOK, P. J. R. 2006. A new snake of the genus Atractus Wagler, 1828 (Reptilia: Squamata: Colubridae) from Kaieteur National Park, Guyana, northeastern South America. Zootaxa, 1378: 19-35 [link]
  • 15.  KOK, P. J. R., SAMBHU, H., ROOPSIND I., LENGLET, G. L. & BOURNE, G. R. 2006. A new species of Colostethus (Anura: Dendrobatidae) with maternal care from Kaieteur National Park, Guyana. Zootaxa, 1238: 35-61 [pdf]
  • 14.  COLE, C. J. & KOK, P. J. R. 2006. A New Species of Gekkonid Lizard (Sphaerodactylinae: Gonatodes) from Guyana, South America. American Museum Novitates, 3524: 1-13 [pdf]
  • 13.  GRANT, T., FROST, D. R., CALDWELL, J. P., GAGLIARDO, R., HADDAD, C. F. B., KOK, P. J. R., MEANS, B. D., NOONAN, B. P, SCHARGEL, W., & WHEELER, W. C. 2006. Phylogenetic systematics of dart-poison frogs and their relatives (Anura: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History, 299: 1-262 [pdf]
  • 12.  KOK, P. J. R., MACCULLOCH, R. D., GAUCHER, P., POELMAN, E. H., BOURNE, G. R., LATHROP, A. & LENGLET, G. L. 2006. A new species of Colostethus (Anura: Dendrobatidae) from French Guiana with a redescription of Colostethus beebei (Noble, 1923) from its type locality. Phyllomedusa, 5(1): 43-66 [pdf]
  • 11.  KOK, P. J. R., BOURNE, G. R., BENJAMIN, P. & LENGLET, G. L. 2006. Stefania evansi (Groete Creek Carrying Frog). Reproduction. Herpetological Review, 37(2): 212-213 [pdf]
  • 10.  KOK, P. J. R., BOURNE, G. R., LAFONTAINE R-M., BENJAMIN, P. & LENGLET, G. L. 2006. Corallus hortulanus (Amazon Tree Boa). Diet. Herpetological Review, 37(2): 230-231



  • 9.  KOK, P. J. R., BOURNE, G. R., SAMBHU, H. & LENGLET, G. L. 2005. Geographic Distribution. Phrynohyas coriaceaHerpetological Review, 36(4): 462-463 [pdf]
  • 8.  KOK, P. J. R. 2005. A New Genus and Species of Gymnophthalmid Lizard (Squamata: Gymnophthalmidae) from Kaieteur National Park, Guyana. Bulletin de l’Institut Royal des Sciences naturelles de Belgique, Biologie, 75: 35-45 [pdf]


  • 7.  KOK, P. J. R., ROZE, J. A., LENGLET, G. L., SAMBHU, H. & ARJOON, D. 2003. Micrurus isozonus (Cope, 1860) (Serpentes, Elapidae): an addition to the herpetofauna of Guyana, with comments on other species of coral snakes from Guyana. Bulletin de l’Institut Royal des Sciences naturelles de Belgique, Biologie, 73: 73-79 [pdf]


  • 6.  KOK, P., JOORIS, R., PERCSY, C. & LENGLET, G. L. 2002. Dangers pour la faune indigène de l’introduction d’espèces animales à des fins ornementales : Rana bedriagae (Amphibia : Anura : Ranidae), un futur cas d’école ? Faune Belge : Etat des Lieux et Tendances Observées (avec une attention particulière pour les espèces exotiques). Bulletin de l’Institut Royal des Sciences naturelles de Belgique, 72(suppl.): 219-221 [pdf]


  • 5.  KOK, P. J. R. 2001. Note sur l’introduction de Rana bedriagae Camerano, 1882 (Anura, Ranidae) en Belgique et ses possibles implications sur la batrachofaune indigène. Les Naturalistes Belges, 82(1): 25-30



  • 4.  KOK, P. J. R. 2000. Addenda to “A survey of the anuran fauna of Montagne Belvédère, county of Saül, French Guiana”. The Herpetological Bulletin, 73: 1 [pdf]
  • 3.  KOK, P. J. R. 2000. A survey of the anuran fauna of Montagne Belvédère, county of Saül, French Guiana: field list with comments on taxonomy and ecology. The British Herpetological Society Bulletin, 71: 6-26 [pdf]


  • 2.  KOK, P. 1998. Ctenophryne geayi (Brown Egg Frog). Geographic Distribution. Herpetological Review, 29(4): 246 [pdf]
  • 1.  KOK, P. 1998. Anolis nitens chrysolepis (Goldenscale Anole). Predation. Herpetological Review, 29(1): 41 [pdf]
Close Menu